All of the flowers now raised here have developed from those______in the forest.A.once they grew B.they grew once C.that once grew D.once grew

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All of the flowers now raised here have developed from those______in the forest.
A.once they grew B.they grew once C.that once grew D.once grew

C
those是定语从句的先行词,在从句中作主语(排除A、B),所以引导词that不可省略(排除D)